Quarterly report pursuant to Section 13 or 15(d)

Cash, Cash Equivalents and Short-Term Investments - Gross Unrealized Gains and Losses (Details)

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Cash, Cash Equivalents and Short-Term Investments - Gross Unrealized Gains and Losses (Details) - USD ($)
$ in Thousands
Sep. 30, 2016
Dec. 31, 2015
Cash equivalents and available-for-sale securities    
Amortized Cost $ 70,350 $ 97,911
Gross Unrealized Gains 19 12
Gross Unrealized Losses (10) (56)
Fair Value 70,359 97,867
U.S. treasury securities    
Cash equivalents and available-for-sale securities    
Amortized Cost 8,301 9,061
Gross Unrealized Gains 6  
Gross Unrealized Losses   (13)
Fair Value 8,307 9,048
U.S. Government-sponsored enterprises securities    
Cash equivalents and available-for-sale securities    
Amortized Cost 16,814 48,643
Gross Unrealized Gains 11 1
Gross Unrealized Losses   (31)
Fair Value 16,825 48,613
Corporate bonds and commercial paper    
Cash equivalents and available-for-sale securities    
Amortized Cost 45,235 40,207
Gross Unrealized Gains 2 11
Gross Unrealized Losses (10) (12)
Fair Value $ 45,227 $ 40,206